It was a matter of manufacturing tolerances, as I've been told. If the sum of the slight errors mostly canceled out, then the 135 worked fine. If they stacked up at one extreme or the other, then there would be focus errors at wider apertures. The UK importer offered a matching service at one time, a good solution. So some unmodified original camera-lens combinations will be fine wide open, and the rest will need to be stopped down somewhat.
Ultimately, Tamron chose to recall all 135mm lenses and issue 100mm lenses in exchange, also replacing the camera frameline mask at no charge. Not all owners availed themselves of this exchange service, so that accounts for the few 00 serial bodies still with 135mm framelines, and the vanishing few 135mm lenses not turned in. All 100 serial bodies will have 100mm framelines, as will most 000 serials too as they have been modified.
I have both types of RF645, having gotten an unmodified older one in hopes of eventually finding the 135 lens, never did.